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Code On Time 8.7.8.0 Unlimited [EXCLUSIVE]

Code On Time 8.7.8.0 Unlimited [EXCLUSIVE]

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Code On Time 8.7.8.0 Unlimited

Subsequent to the ancestral distribution, the ancestors of the Japanese population migrated towards Hokkaido around the tMRCA of the Japanese black bears (0.99 Ma) (Fig 3 and Fig 4). According to the transition model with the TEMPORAL program, the population size of the ancestral population was decreasing in a linear fashion. The ancestral population grew linearly until about 2.6 Ma. The size of the population dropped sharply around the tMRCA of Japanese black bears. Since the most recent common ancestor (MRCA) of the Japanese population is younger than the divergence time of the western subspecies in European black bears (2.39 Ma), this population presumably expanded from a local source population, probably around the 1.3 Ma. Thus, we can speculate that the Japanese black bears gradually expanded from the Japanese archipelago and today inhabit a wide area of northern Japan. Such a topographic model (i.e., the subspecies expansion) has been considered in other studies [ 68 – 71 ], including black bears in North America [ 72 ] and the Iberian peninsula [ 73 ].

The Asian black bears in Hokkaido, as well as the Japanese population, are distributed in the northern part of Honshu and the west coast of the island, and are genetically differentiated from the populations in the southern part of Honshu, Kyushu, and the Korean Peninsula. The current distribution of black bears in Hokkaido is 0.12 to 0.94 times larger than that of the continental population (Table 2). The distribution of brown bears is also shown in Fig 2. Brown bears are distributed in Hokkaido [ 73 – 75 ], and their range is partially overlapping with that of Asian black bears. The historical records of Hokkaido showed that brown bears have been distributed in Hokkaido since about the 17th century [ 76 – 78 ]. Based on the stable isotope analysis, the recent migration event of the brown bears from the mainland was observed to be about 100 years ago [ 73, 79 ]. In the native range of European black bears, there is currently a division into two clusters: one of European black bears and one of continental black bears [ 29, 30 ]. It was suggested that the current distribution of the European black bears is due to the geographical separation of populations between the eastern Mediterranean area and west Europe during the LGM. Our topological model of the ML tree and the MJ network also supported this hypothesis. The European black bears and the Asian black bears are clustered in the separate subclades.

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